The game begins in 1911, where
John Marston (
Rob Wiethoff), a former
outlaw, is taken from his family by the
Bureau of Investigation. They state Marston will be granted amnesty if he brings the remaining members of his old gang to justice. Marston agrees and sets off to track down Bill Williamson (Steve J. Palmer), who now runs his own gang. Marston confronts Williamson at his stronghold, Fort Mercer, only to be shot and left for dead. A local rancher, Bonnie MacFarlane (Kimberly Irion), finds him critically wounded and takes him to her ranch for recovery. Several days later, Marston begins repaying the MacFarlanes for their help in the form of odd-jobs around the ranch. During this time he works with a
U.S. Marshal, Leigh Johnson (
Anthony De Longis) and his deputies, the dim-witted Jonah and sarcastic Eli; Nigel West Dickens (
Don Creech), a con artist selling
patent medicines; Seth Briars (Kevin Glikmann), a treasure hunter and grave robber who prefers the company of the dead to that of the living; and an unreliable alcoholic arms dealer known only as Irish (K. Harrison Sweeney). Marston performs various tasks and favours in exchange for their help with organizing an attack on Fort Mercer. Marston and his group then breach the fort and defeat Williamson's gang only to find that Williamson had already fled to Mexico to seek help from Javier Escuella (Antonio Jaramillo), another member of Marston's former gang.
In Mexico, Marston initially sides with Mexican Army Colonel Agustín Allende (Gary Carlos Cervantes) and his right-hand man Captain Vincente de Santa (
Hector Luis Bustamante), to help put down a rebellion in return for Allende delivering Escuella and Williamson to him. However, Allende betrays Marston, who is then forced to side with the rebels. With the initial help of elderly gunslinger Landon Ricketts (
Ross Hagen), Marston, along with rebel leader Abraham Reyes (Josh Segarra), and Luisa (Francesca Galeas), one of Reyes' many lovers, eventually turns the tide in favour of the rebels, killing de Santa and launching an assault on the El Presidio fortress, where they find Escuella, who attempts to bargain for his life by revealing that Williamson has been under Allende's protection. Escuella manages to escape, with Marston giving chase, who is then given the option to either kill or capture Escuella alive and turn him over to the Bureau. Either way, agents Edgar Ross (Jim Bentley) and Archer Fordham (David Wilson Barnes), who meet with Marston at the USA/Mexico border, refuse to let Marston return to his family, stating that after killing Williamson, he must meet with them again in Blackwater and assist in hunting down Dutch van der Linde (Benjamin Byron Davis), the leader of Marston's former gang. Marston and Reyes then take the fight to Allende, where Luisa is killed during the attack. As Allende's palace is stormed by the rebels, both he and Williamson flee, who are then apprehended and executed after being chased a short distance. Reyes then takes charge of the local government and plans to advance on the capital, while Marston leaves to meet with the Bureau in Blackwater.
Once there, Marston learns that Dutch has been forming his own gang with sympathizers from the local Indian reservation, and joins Ross, Fordham and a group of U.S. soldiers in an attack on Dutch's hideout. During the assault, Marston chases Dutch up to a cliff overlooking the river and mountains. Dutch ultimately chooses to commit suicide, backing off the cliff and falling to his death, but not before warning Marston that the Bureau will 'just find another monster' to justify their pay. Marston is then released from his deal with Ross and goes back to his ranch where his wife Abigail (Sophie Marzocchi) and his son Jack (
Josh Blaylock) are waiting. Marston devotes his time afterward tending to the ranch, having sworn to keep himself and his family away from the outlaw lifestyle forever. He is then suddenly forced to repel a surprise attack from a combined force of soldiers, lawmen and government agents led by Ross. Marston is able to fend off several waves of attackers and get his family to safety but stays behind to hold back the rest. Despite a valiant last stand, he dies after being shot numerous times by Ross and his men. He is then buried by his grieving family on the hill overlooking the ranch.
The game then moves three years ahead to 1914, where Marston's son Jack stands over the graves of both his father and his now deceased mother. He leaves home to search for Ross, who retired the year before, and eventually finds the former agent hunting on a riverbank in Mexico. The two duel, with Jack emerging the victor. The fate of John Marston's companions are later summarized: Marshal Leigh Johnson retired from Armadillo to be replaced by Jonah, and then moved as far as possible from the town; Abraham Reyes, though promising to lead Mexico into an age of freedom, was consumed by power and became a tyrant; Bonnie MacFarlane eventually married; Irish shot himself dead in an outhouse when his gun accidentally discharged; Seth Briars eventually found the treasure he was looking for and became rich; and finally, Landon Ricketts died quietly in his sleep.
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